Career

1965 M.A. (Physics), Cambridge University.

1966 Diploma Adv. Physics, Manchester University.

1968 PhD (Biophysics), Manchester University.

1969-94 MRC Human Genetics Unit (Edinburgh): Appointed to Scientific Staff

1973 Research Fellow, Dept Anatomy, Harvard University, Boston, USA (1 year).

1975 Appointed MRC tenured scientist

1980 Visiting Professor, Applied Math. Dept. Weizmann Inst. Israel (4m).

1982-3 Seconded to the Rayner "Efficiency" Office (Whitehall) and MRC HQ to
undertake a scrutiny of MRC Laboratory Management.

1985 Promoted to Senior Scientist (Reader grade).

1994 Edinburgh University Senior lecturer

1996 Reader in Developmental Anatomy

2002 Personal chair in Development and Bioinformatics

2007 Retired with Emeritus Professor and Senior Fellow status

2007-9 Honorary member of the Computational Biology Group (WIMM, Oxford University)

2009- Visiting member, Dept. Physiology, Anatomy & Genetics, Oxford University