Career
1965 M.A. (Physics), Cambridge University.
1966 Diploma Adv. Physics, Manchester University.
1968 PhD (Biophysics), Manchester University.
1969-94 MRC Human Genetics Unit (Edinburgh): Appointed to Scientific Staff
1973 Research Fellow, Dept Anatomy, Harvard University, Boston, USA (1 year).
1975 Appointed MRC tenured scientist
1980 Visiting Professor, Applied Math. Dept. Weizmann Inst. Israel (4m).
1982-3
Seconded to the Rayner
"Efficiency" Office (Whitehall)
and MRC HQ to
undertake a scrutiny of MRC Laboratory
Management.
1985 Promoted to Senior Scientist (Reader grade).
1994 Edinburgh University Senior lecturer
1996 Reader in Developmental Anatomy
2002 Personal chair in Development and Bioinformatics
2007 Retired with Emeritus Professor and Senior Fellow status
2007-9 Honorary member of the Computational Biology Group (WIMM, Oxford University)
2009- Visiting member, Dept. Physiology, Anatomy & Genetics, Oxford University